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<aff id="hhMI">Department of Molecular, Cellular, and Developmental
Biology, Howard Hughes Medical Institution, University of Colorado,
Boulder, Colorado 80303-0347</aff>
<author-notes>
<fn id="FN150"><p>Corresponding author. E-mail
address: <email>mhan@colorado.edu</email>.</p></fn>
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<notes>
<p>Helpful comments from two anonymous referees on earlier
version of the paper are gratefully acknowledged. Email:
<email><italic>kuse@udel.edu</italic></email></p>
</notes>
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Journal Archiving and Interchange Tag Library NISO JATS Draft Version 1.1d2
Version of December 2014